How did written punctuation originate?

It wasn’t until the end of the fifteenth century that the Italian printer Aldus Manutius introduced the system of markings we call punctuation. The proper use of punctuation marks is a learned skill that has eluded even great writers ever since. Mark Twain once filled the last page of a manuscript with all the various symbols of punctuation and instructed his editor to disperse them within the story as he saw fit.

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